#NEXUS
[MacClade 3.01 registered to Equator, UCMP]

BEGIN DATA;
DIMENSIONS  NTAX=27 NCHAR=113;

[!This is the final revised morphological matrix for land plants (LP-MORPH),
corrected 12/93,  per comments of co-authors.

**used in the Ann .Mo. Bot. 81: 451-483 (Mishler et al. 1994) green plant synthesis paper.

Characters are re-numbered as they are used in our text, but you can see Karen Renzaglia's old
numbers (from the sperm data set, Garbary et al. 1993, Pl. Syst. & Evol.,
which see for more details on characters), or reference to other literature
(M&C = Mishler and Churchill, 1985, Cladistics 1:305-328) as footnotes to the characters.]
FORMAT MISSING=? GAP=-  SYMBOLS= " 0 1 2 3 4 5 6";
[OPTIONS  MSTAXA=UNCERTAIN ;]

CHARLABELS
	[1]	apical_cell_in_antheridia	[2]	div._pat._in_young_antheridia	[3]	endogenous_antheridia	[4]	antheridial_stalk	[5]	operculum_cells
	[6]	sperm_in_pollen_tube	[7]	number_of_sperm_per_male_struc.	[8]	nascent_spermatids	[9]	diagonal_spindle_in_final_mitot	[10]	replication_of_the_centrioles
	[11]	time_of_origin_of_centrioles	[12]	basal_bodies_and_flagella	[13]	bicentrioles	[14]	basal_body_position	[15]	proximal_extension_A
	[16]	proximal_extension_B	[17]	stellate_transition	[18]	connect._fibers_betw._basal_bod	[19]	basal_body_structure	[20]	BB_staggering_assoc._w._MT_grow
	[21]	regression_of_LS	[22]	'LS/AM_elongation_longit.'	[23]	spline_aperture	[24]	sline_aperture_location	[25]	position_of_developing_MLS
	[26]	plaque_stratified_betw._blephar	[27]	'spline/LS_orientation'	[28]	posterior_notch_to_LS	[29]	LS_position	[30]	stray_spline_MT
	[31]	accessory_band_of_MTs	[32]	maturational_elongation_of_AM	[33]	spline_shank	[34]	osmiophilic_crest	[35]	anterior_osmiophilic_crest
	[36]	changes_in_BBs_at_mat.	[37]	matrix_around_BBs	[38]	posterior_of_the_stellate_patt.	[39]	flagellar_scales	[40]	late_blepharoplast_w._transient
	[41]	direction_of_flagellar_emergenc	[42]	nuclear_shape_at_maturity	[43]	nuclear_posterior_shape	[44]	median_constriction	[45]	splin_attached_to_nucleus
	[46]	spline_growth_assoc._w._nucl._s	[47]	direction_of_nuclear_compaction	[48]	condensed_chromatin_strands	[49]	diverticulum_during_shaping	[50]	number_of_gyres_of_nucleus
	[51]	dense_body_in_ant._mitochondrio	[52]	mit._assoc._w._plastids_in_sper	[53]	mit._assoc._w._plastids_in_sper	[54]	specialized_ant._mito.	[55]	specialized_posterior_mito.
	[56]	additional_mito._in_ant._of_cel	[57]	origin_of_AM	[58]	osmophilic_material_underneath	[59]	change_from_cristae_sacs_to_baf	[60]	monoplastidic_sperm
	[61]	plastid_determines_division_pol	[62]	starch_grains_in_single_plastid	[63]	sperm_plastid_contacting_nucleu	[64]	fibrillenscheide	[65]	cytoplasmic_loss
	[66]	embryo	[67]	cuticle	[68]	lunularic_acid	[69]	elaters	[70]	oil_bodies
	[71]	'D-Methionine'	[72]	stomates	[73]	vert_div_of_zygote	[74]	xylem	[75]	phloem
	[76]	perine_layer_on_spores	[77]	'aerial_sporophyte_axis_(persist'	[78]	columella	[79]	Multicellular_rhizoids	[80]	leaves_on_gam.
	[81]	independent_sporophyte	[82]	branched_sporophyte	[83]	tracheids	[84]	lignin	[85]	'long_exserted_seta_(of_liv._typ'
	[86]	oil_body_cells	[87]	spore_mother_cells_lobed	[88]	'capsule_2-4_valved'	[89]	'caps_wall_cells_w/_trans_thick'	[90]	thick_caps._wall
	[91]	elongate_antheridia	[92]	aerial_calyptra	[93]	paraphyses	[94]	costate_leaves	[95]	peristome
	[96]	operculum	[97]	cylindical_sprogen_layer	[98]	transfer_cells_on_gam_side	[99]	transfer_cells_on_spor.__side	[100]	seed
	[101]	megaphyll	[102]	microphyll	[103]	'lateral,_broad_sporangia'	[104]	exarch_maturation_of_xylem	[105]	cp_DNA_inversion
	[106]	sporangia_borne_on_leaves	[107]	trichomes	[108]	flavonoids	[109]	retention_of_zygote	[110]	sheathed_hairs
	[111]	polyphenolics_induced_by_sex	[112]	parenchyma	[113]	beaked_muscilage_papillae
	;
STATELABELS
	1  absent present,
	2  'four-celled' 'two-celled',
	3  absent present,
	4  absent present,
	5  absent present,
	6  absent present,
	7  1000+ '100-1000' '16-24' 2,
	8  paired not_paired,
	9  absent present,
	10  present absent,
	11  always_present sperm._mother_cells sperm._moth._cell_progenitor earlier,
	12  two more_than_two,
	13  present absent,
	14  right_angles 'side-by-side' 'staggered_ant-post' staggered_continous,
	15  absent long short,
	16  'ventral_-_dorsal' ventral,
	17  present absent,
	18  present absent fine_filaments_w._centrin,
	19  monomorphic dimorphic,
	20  absent present,
	21  absent complete partial,
	22  parallel perpendicular,
	23  absent present,
	24  left_of_center right_of_center,
	25  adjacent_to_BBs beneath_BBs,
	26  absent present,
	27  90_deg. 45_deg.,
	28  absent present,
	29  under_all_BBs under_ABB_only under_some_BBs,
	30  absent present develops_late,
	31  absent present,
	32  absent posterior,
	33  wide less_than_4_tubules,
	34  absent present,
	35  absent present,
	36  absent dense_material_at_tip BB_cartwheel_w._plug BB_triplets_impreg._w._matrixma,
	37  homogenous mottled,
	38  extracell._or_partly entirely_intracell.,
	39  present absent,
	40  yes no,
	41  toward_side toward_rear toward_anterior,
	42  ovoid elongate,
	43  not_expanded expanded,
	44  absent present,
	45  yes detached_at_maturity never_attached,
	46  absent present,
	47  outer_shell anterior_to_posterior at_equal_rates_along_nucl. general_increase_in_density,
	48  'spaghetti-like' perpendicular_to_spline 'spiral-central_strand' general_compaction spikes irregular_plates solid_mass_from_ant._tip,
	49  absent present,
	50  not_coiled '0.5-3' greater_than_3,
	51  absent present,
	52  absent present,
	53  absent present,
	54  present absent,
	55  present absent,
	56  absent row_of_mito._behind_AM numerous_unspecialized,
	57  fusion elongation,
	58  absent present,
	59  absent present,
	60  present absent,
	61  'present_-_at_poles' 'present_-_asymmetrical' absent,
	62  more_than_one one,
	63  absent present,
	64  absent present,
	65  absent partial 'complete_(or_tiny_remn.)',
	66  absent present,
	67  absent present,
	68  absent present,
	69  absent present,
	70  absent present,
	71  not_recognize recognize,
	72  absent present,
	73  absent present,
	74  absent present,
	75  absent present,
	76  absent present,
	77  absent present,
	78  absent present,
	79  absent present,
	80  absent 'present_(of_moss_type)' 'present_(of_junger._type)',
	81  absent present,
	82  absent present,
	83  absent present,
	84  absent present,
	85  absent present,
	86  absent present,
	87  absent present,
	88  absent present,
	89  absent present,
	90  'absent_(unistrat.)' 'present_(2-10_strat.)',
	91  absent present,
	92  absent present,
	93  absent present,
	94  absent present,
	95  absent present,
	96  absent present,
	97  absent present,
	98  absent present,
	99  absent present,
	100  absent present,
	101  absent present,
	102  absent present,
	103  absent present,
	104  absent present,
	105  absent present,
	106  absent present,
	107  absent present,
	108  absent present,
	109  absent present,
	110  absent present,
	111  absent present,
	112  absent present,
	113  absent present,
;

MATRIX
					 [----+--10|----+--20|----+--30|----+--40|----+--50|----+--60|----+--70|----+--80|----+--90|----+-100|----+-110|---]
CHARA            ?????0??0030?00?020???0??0???00?0000000?11000000010??1?1?011201020000000000000000000000000?0000000000000?00100000
NITELL           ?????0??0030?00?020???0?000??00?0000000?11000000010001???0?1201020000000000000000000000000?0000000000000?00100000
COLEOC_PUL       ?????0???000?00?0?0?000?000??0000000000?00?00???000000?1??0000?010000000000000000000000000?0000000000000?00011100
COLEOC_ORB       ?????0??1000?00?0?0?000?000??0000000000?00?00???000000?1??0000?010000000000000000000000000?0000001000000?00011110
PHAEOCEROS       0011000001101111100?200?101000010003001?1101001101111000011011102110001110000100000000000000000001000000?00010110
NOTOTHYLAS       0011000001101111100?200?101000010003001?1101001101111000011011102110001110000100000000000000000001000000?00010110
MARCHANTIA       000100000110121001112011101100000003001?1100001011001000001010112111110000000000000001000000000001100000000110110
SPHAEROCARPOS    000100000110121001112011101100001003001?1100001011001000001010??211111000000000000000{01}000000000001100000?00110110
PELLIA           010100000110121001112011101000000003001?1100101011001000001010112111110000000000000010111100000000000000?00110110
BLASIA           0?0100000110121001112011101100000003001?1100001011001000001010112111110000000000000010111100000001100000?00110110
JUNGERMANNIA     010100000110121001112011101000000003001?1100101011001000001010112111110000000002000010111100000000100000?00110110
HAPLOMITRIUM     010100000110121?01112010101000?00?03001?11000015?10010000010?1??2111110000000000000010110000000001100000?00110110
TREUBIA          01010000011012100111?010101000?00?03?01?11?0001???0??0000??0?0??211111000000000000001011110000000??00000?00110110
SPHAGNUM         110100011110121011111011101012010003001?1100012201011000001001002110001100011111000000000001000100000000?00010110
ANDREAEA         1101100111101210?1112011101010010003001?1100012201011000001000002110001001011111000000000011110000100000?00010111
POLYTRICHUM      110110011110121001112011101011011003001?1100012201011000001000002110001101111111000000000011111110100000?00010110
HYPNUM           110110011110121001112011101011010003001?1100012211011000001000002110001101111111000000000011111111100000000110110
TAKAKIA          11011001111012100111?01110101??10?03?01?1100?????10?10000?1000??2110001001??11?10000000000110?0000100000?00110111
LYCOPODIELLA     0?00001?1110140?020?010?1010000?0?00101?10?000??00001010?000?0001110001101111000111100000000000001100111000110110
LYCOPODIUM       0?00001011101221020?010?101000010000101?10000?3310000010?0100010{12}110001101111000111100000000000001100111000110110
SELAGINELLA      0??0?01??110?2210?0?200?101000010000101?110000?401001000?010?000211000110111100011110000000000000??00111000110110
EQUISETUM        0?0000111111030?0100010?1110201001100011111000330100001110012?00111000110111100011110000000000000??00?00110110110
PTERIDIUM        0?000021112103210?00010?111020100112?011111000330200001110012?001110001111111000111100000000000001101000111110110
MARSILEA         0??0?0211121030?0100010?1110200000020011110000160200001010012?00111000111111100011110000000000000??01000?11110110
OSMUNDA          0?000021112103210?00010?1110201001110011111000330100001110012?001110001111111000111100000000000001101000111110110
GINKGO           ?????13?11?1030?0100010?1?1020000110011020?02???00000112?0012?0001100011011?100011110000000000000??11000111110110
ZAMIA            ?????13?11?1030?0100010?1?1020000110011020?02???00000112?0012?0001100011011?100011110000000000000??11000111110110
;
END;

BEGIN ASSUMPTIONS;
	OPTIONS  DEFTYPE=unord PolyTcount=MINSTEPS ;

	TYPESET * order_two  = unord: 1-6 8-49 51-113, ord: 7 50;

	WTSET * CURRENT  = 1: 1-113;

	EXSET  sperm_only  =  1-65;
	EXSET  general_morph  =  66-113;

END;
begin paup;
	constraint moss+trach =  (1,2,3,4,5,6,7,8,9,10,11,12,13,(14,15,16,17,18,19,20,21,22,23,24,25,26,27));
	constraint horn+moss+trach = (1,2,3,4,7,8,9,10,11,12,13,(5,6,14,15,16,17,18,19,20,21,22,23,24,25,26,27));
	constraint bryos = (1,2,3,4,(19,20,21,22,23,24,25,26,27,(5,6,7,8,9,10,11,12,13,14,15,16,17,18)));
	
end;

BEGIN NOTES;
	TEXT  CHARACTER=1 TEXT= 'KAREN''S_#2';
	TEXT  CHARACTER=2 TEXT= 'KAREN''S_#3';
	TEXT  CHARACTER=3 TEXT= 'KAREN''S_#5';
	TEXT  CHARACTER=4 TEXT= 'KAREN''S_#6';
	TEXT  CHARACTER=5 TEXT= 'KAREN''S_#7';
	TEXT  CHARACTER=6 TEXT= 'KAREN''S_#8';
	TEXT  CHARACTER=7 TEXT= 'KAREN''S_#10__!ORD';
	TEXT  CHARACTER=8 TEXT= 'KAREN''S_#12';
	TEXT  CHARACTER=9 TEXT= 'KAREN''S_#14';
	TEXT  TAXON=20 TEXT= 'Lycopodium_obscurum_added_from_Karen''s_notes_sent_3/1/93_--_note_that_the_old_Lycop_is_now_Lycopodiella_!';
	TEXT  CHARACTER=10 TEXT= 'KAREN''S_#15';
	TEXT  CHARACTER=11 TEXT= 'KAREN''S_#16';
	TEXT  CHARACTER=12 TEXT= 'KAREN''S_#17	';
	TEXT  CHARACTER=13 TEXT= 'KAREN''S_#18';
	TEXT  CHARACTER=14 TEXT= 'KAREN''S_#19		';
	TEXT  CHARACTER=15 TEXT= 'KAREN''S_#20';
	TEXT  CHARACTER=16 TEXT= 'KAREN''S_#21';
	TEXT  CHARACTER=17 TEXT= 'KAREN''S_#22';
	TEXT  CHARACTER=18 TEXT= 'KAREN''S_#24';
	TEXT  CHARACTER=19 TEXT= 'KAREN''S_#25';
	TEXT  CHARACTER=20 TEXT= 'KAREN''S_#26';
	TEXT  CHARACTER=21 TEXT= 'KAREN''S_#27';
	TEXT  CHARACTER=22 TEXT= 'KAREN''S_#28';
	TEXT  CHARACTER=23 TEXT= 'KAREN''S_#30';
	TEXT  CHARACTER=24 TEXT= 'KAREN''S_#31';
	TEXT  CHARACTER=25 TEXT= 'KAREN''S_#32';
	TEXT  CHARACTER=26 TEXT= 'KAREN''S_#34';
	TEXT  CHARACTER=27 TEXT= 'KAREN''S_#35';
	TEXT  CHARACTER=28 TEXT= 'KAREN''S_#37';
	TEXT  CHARACTER=29 TEXT= 'KAREN''S_#38';
	TEXT  CHARACTER=30 TEXT= 'KAREN''S_#39';
	TEXT  CHARACTER=31 TEXT= 'KAREN''S_#40';
	TEXT  CHARACTER=32 TEXT= 'KAREN''S_#44';
	TEXT  CHARACTER=33 TEXT= 'KAREN''S_#47';
	TEXT  CHARACTER=34 TEXT= 'KAREN''S_#49';
	TEXT  CHARACTER=35 TEXT= 'KAREN''S_#50';
	TEXT  CHARACTER=36 TEXT= 'KAREN''S_#51';
	TEXT  CHARACTER=37 TEXT= 'KAREN''S_#52';
	TEXT  CHARACTER=38 TEXT= 'KAREN''S_#53';
	TEXT  CHARACTER=39 TEXT= 'KAREN''S_#54';
	TEXT  CHARACTER=40 TEXT= 'KAREN''S_#55';
	TEXT  CHARACTER=41 TEXT= 'KAREN''S_#57';
	TEXT  CHARACTER=42 TEXT= 'KAREN''S_#58';
	TEXT  CHARACTER=43 TEXT= 'KAREN''S_#59';
	TEXT  CHARACTER=44 TEXT= 'KAREN''S_#61';
	TEXT  CHARACTER=45 TEXT= 'KAREN''S_#63';
	TEXT  CHARACTER=46 TEXT= 'KAREN''S_#64';
	TEXT  CHARACTER=47 TEXT= 'KAREN''S_#65';
	TEXT  CHARACTER=48 TEXT= 'KAREN''S_#66';
	TEXT  CHARACTER=49 TEXT= 'KAREN''S_#67	';
	TEXT  CHARACTER=50 TEXT= 'KAREN''S_#69__ORD!';
	TEXT  CHARACTER=51 TEXT= 'KAREN''S_#70';
	TEXT  CHARACTER=52 TEXT= 'KAREN''S_#71';
	TEXT  CHARACTER=53 TEXT= 'KAREN''S_#72';
	TEXT  CHARACTER=54 TEXT= 'KAREN''S_#74';
	TEXT  CHARACTER=55 TEXT= 'new_character_added_by_Karen_3/1/93';
	TEXT  CHARACTER=56 TEXT= 'KAREN''S_#75';
	TEXT  CHARACTER=57 TEXT= 'KAREN''S_#76';
	TEXT  CHARACTER=58 TEXT= 'KAREN''S_#77';
	TEXT  CHARACTER=59 TEXT= 'KAREN''S_#79';
	TEXT  CHARACTER=60 TEXT= 'KAREN''S_#80';
	TEXT  CHARACTER=61 TEXT= 'KAREN''S_#81';
	TEXT  CHARACTER=62 TEXT= 'KAREN''S_#83';
	TEXT  CHARACTER=63 TEXT= 'KAREN''S_#84';
	TEXT  CHARACTER=64 TEXT= 'KAREN''S_#86';
	TEXT  CHARACTER=65 TEXT= 'KAREN''S_#89';
	TEXT  CHARACTER=77 TEXT= 'of_moss/tracheophyte_type';
	TEXT  CHARACTER=80 TEXT= unord.;
	TEXT  CHARACTER=85 TEXT= 'of_the_liverwort_type_(non-homologous_to_those_of_the_moss/tracheophyte_lineage)__--__M&C_char._#9';
	TEXT  CHARACTER=88 TEXT= 'on_a_regular_basis_(Andreaea_considered_non-homologous)_--_M&C_char._#14';
	TEXT  CHARACTER=108 TEXT= see_Markham_in_Zinsmeister_&_Mues;
	TEXT  CHARACTER=90 TEXT= 'caps._wall_2-10_cells_thick_--__M&C_char._#19_--_thick_caps._walls_in_horn.,_mosses,_and_tracheo._considered_non-nomologous';
	TEXT  CHARACTER=89 TEXT= M&C_char._#18;
	TEXT  CHARACTER=86 TEXT= M&C_char._#10;
	TEXT  CHARACTER=87 TEXT= M&C_char._#13;
	TEXT  TAXON=18 CHARACTER=80 TEXT= considered_homologous_with_mosses;
	TEXT  TAXON=15 CHARACTER=94 TEXT= assuming_loss_in_some_spp.;
	TEXT  TAXON=17 CHARACTER=94 TEXT= assuming_loss_in_some_spp.;
	TEXT  TAXON=15 CHARACTER=74 TEXT= possible_homology_in_Andreobryum_?;
	TEXT  CHARACTER=82 TEXT= with_multiple_sporangia;
	TEXT  CHARACTER=109 TEXT= through_sporogenesis;
	TEXT  TAXON=15 CHARACTER=72 TEXT= fide_Murray_1988;
	TEXT  TAXON=15 CHARACTER=93 TEXT= fide_Murray_1988;
	TEXT  TAXON=15 CHARACTER=113 TEXT= 'in_Andreaobryum_only_(fide_Murray_1988),_Andreaea_has_normal_moss-like_muscilage_papillae';
	TEXT  CHARACTER=113 TEXT= homology_accepted_from__Murray_1988;
	TEXT  CHARACTER=106 TEXT= Bremer_char._#_64;
	TEXT  CHARACTER=107 TEXT= Bremer_#_67;
	TEXT  CHARACTER=105 TEXT= from_Raubeson_and_Jansen_1992;
	TEXT  CHARACTER=103 TEXT= Crane;
	TEXT  CHARACTER=104 TEXT= Crane;
	TEXT  CHARACTER=102 TEXT= Crane;
	TEXT  TAXON=4 CHARACTER=9 TEXT= not_sure_of_homology_with_mosses_+_tracheophytes;
	TEXT  CHARACTER=74 TEXT= 'N.B.,_homology_between_mosses_and_tracheophytes_not_agreed_to_by_all_co-authors';
	TEXT  CHARACTER=75 TEXT= 'N.B.,_homology_between_mosses_and_tracheophytes_not_agreed_to_by_all_co-authors';
	TEXT  TAXON=8 CHARACTER=86 TEXT= 'polymorphic_in_order;__Riella_has_differentiated_cells';
	TEXT  TAXON=18 CHARACTER=94 TEXT= 'state_difficult_to_determine_--_maybe_all_costa!';
	TEXT  TAXON=22 CHARACTER=102 TEXT= not_sure_if_homologous_with_lycophytes;
END;

BEGIN MACCLADE;
v 3.0
-1322664995
1100&/0
0
0
END;
